@grimalkina Hyper-specialization requires a societal substrate to support it. To early and the web is too fragile, the specialist needs the support of the generalist to fill the gaps.
Over time, if the society grows large enough, the gaps between the nodes in the mesh narrow as normal randomization within standard distribution spreads. This allows more specialization within the same structure without requiring changes to the constraints of the overall system.
The specialist can be equally well supported in either case, though that support must be there or they will collapse. Project the singular case upon an entire society where the ratio of specialist to generalist is skewed and my feeling is that would ultimately lead to collapse
Take what you would from that as metaphor for specialized or exceptional workers in a typical corporate environment.